/* BRAND-001D scientific public register.
 * BRAND-001A owns the tokens and BRAND-001B owns the shell. This file only
 * gives Taxonomy, Catalogue, Distribution and Data distinct work surfaces. */

:root {
  --science-bg: var(--shell-bg);
  --science-surface: var(--shell-surface);
  --science-surface-2: var(--shell-surface-2);
  --science-ink: var(--shell-text);
  --science-muted: var(--shell-muted);
  --science-line: var(--shell-border);
  --science-link: var(--shell-link);
  --science-accent: var(--shell-accent);
  --science-moss: var(--brand-moss-700);
  --science-indigo: var(--brand-indigo-900);
  --science-danger: var(--brand-danger, #a43b2d);
  --science-max: 1240px;
  --science-reading: 760px;
  /* Stable public-catalog aliases retained for the released contrast contract. */
  --bg: var(--science-bg);
  --ink: var(--science-ink);
  --muted: var(--science-muted);
  --amber: var(--science-accent);
}

[data-theme="dark"] {
  /* Neutral 400 falls below AA on the elevated scientific surface. */
  --science-muted: var(--brand-neutral-200);
  --science-moss: var(--brand-paramo-500);
  --science-indigo: var(--brand-indigo-400);
  --science-danger: #f1a093;
}

body[data-page="taxonomy"],
body[data-page="catalog"],
body[data-page="map"],
body[data-page="data"] {
  margin: 0;
  background: var(--science-bg);
  color: var(--science-ink);
  font: 400 1rem/1.58 var(--brand-font-body);
  -webkit-font-smoothing: antialiased;
}

body[data-page="taxonomy"] *,
body[data-page="catalog"] *,
body[data-page="map"] *,
body[data-page="data"] * {
  box-sizing: border-box;
}

body[data-page="taxonomy"] main,
body[data-page="catalog"] main,
body[data-page="map"] main,
body[data-page="data"] main {
  width: min(calc(100% - 40px), var(--science-max));
  margin: 0 auto;
  padding-top: var(--shell-height);
}

body[data-page="taxonomy"] :is(button, input, select),
body[data-page="catalog"] :is(button, input, select),
body[data-page="map"] :is(button, input, select),
body[data-page="data"] :is(button, input, select) {
  font: inherit;
}

body[data-page="taxonomy"] :is(a, button, input, select):focus-visible,
body[data-page="catalog"] :is(a, button, input, select):focus-visible,
body[data-page="map"] :is(a, button, input, select):focus-visible,
body[data-page="data"] :is(a, button, input, select):focus-visible {
  outline: 3px solid var(--science-accent);
  outline-offset: 3px;
}

.eyebrow,
.science-kicker,
.metric-label,
.metric-caption,
.rank,
.taxonomy-path,
.capability-state {
  margin: 0 0 10px;
  color: var(--science-link);
  font: 700 var(--brand-text-xs)/1.25 var(--brand-font-ui);
  letter-spacing: .105em;
  text-transform: uppercase;
}

.catalog-hero,
.map-hero,
.science-hero {
  padding: clamp(56px, 8vw, 104px) 0 clamp(40px, 6vw, 72px);
  border-bottom: 1px solid var(--science-line);
}

.catalog-hero,
.science-hero--split {
  display: grid;
  grid-template-columns: minmax(0, 1.35fr) minmax(300px, .65fr);
  gap: clamp(32px, 6vw, 88px);
  align-items: end;
}

.map-hero,
.science-hero__copy {
  max-width: 880px;
}

.catalog-hero h1,
.map-hero h1,
.science-hero h1 {
  max-width: 900px;
  margin: 0;
  font: 700 clamp(2.55rem, 5.5vw, 4.85rem)/.96 var(--brand-font-display);
  letter-spacing: -.052em;
  text-wrap: balance;
}

.lead,
.science-lead {
  max-width: var(--science-reading);
  margin: 22px 0 0;
  color: var(--science-muted);
  font-size: clamp(1.05rem, 1.8vw, 1.28rem);
  line-height: 1.5;
}

.science-boundary {
  align-self: stretch;
  padding: 24px;
  border: 1px solid var(--science-line);
  background: var(--science-surface);
}

.science-boundary h2 {
  margin: 0 0 12px;
  font: 700 clamp(1.55rem, 3vw, 2.1rem)/1.05 var(--brand-font-display);
  letter-spacing: -.035em;
}

.science-boundary p:last-child { margin-bottom: 0; color: var(--science-muted); }

.snapshot-card,
.snapshot-context {
  padding: 24px;
  border: 1px solid var(--science-line);
  background: var(--science-surface);
}

.snapshot-context { margin: 32px 0 16px; }

.snapshot-card .metric-value {
  display: block;
  margin: 4px 0;
  font: 700 clamp(2.5rem, 4vw, 4rem)/1 var(--brand-font-display);
  font-variant-numeric: tabular-nums;
}

.snapshot-meta {
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: 0 18px;
  margin: 20px 0 0;
}

.snapshot-context .snapshot-meta {
  grid-template-columns: repeat(5, minmax(0, 1fr));
}

.snapshot-meta div,
.taxonomy-grid div {
  padding: 10px 0;
  border-top: 1px solid var(--science-line);
}

dt { color: var(--science-muted); }
dd { margin: 0; color: var(--science-ink); font-weight: 700; overflow-wrap: anywhere; }

.snapshot-card p,
.snapshot-context p { color: var(--science-muted); }

.catalog-shell,
.map-search,
.table-section,
.taxonomy-workbench,
.science-panel {
  margin-top: 16px;
  padding: clamp(24px, 4vw, 48px);
  border: 1px solid var(--science-line);
  background: var(--science-surface);
}

.catalog-shell { margin-top: 32px; }

.section-heading,
.science-section-heading {
  display: flex;
  align-items: end;
  justify-content: space-between;
  gap: 32px;
}

.section-heading h2,
.science-section-heading h2,
.source-note h2,
.map-copy h2,
.science-panel h2 {
  margin: 0;
  font: 700 clamp(1.8rem, 3.8vw, 3rem)/1 var(--brand-font-display);
  letter-spacing: -.042em;
  text-wrap: balance;
}

.secondary-link,
.text-button,
.source-links a,
.detail-actions a,
.science-link {
  display: inline-flex;
  align-items: center;
  min-height: 44px;
  color: var(--science-link);
  font: 700 var(--brand-text-sm)/1.2 var(--brand-font-ui);
  text-underline-offset: .22em;
}

.filters {
  display: grid;
  grid-template-columns: 2fr 1fr 1fr auto;
  gap: 16px;
  align-items: end;
  margin: 30px 0 22px;
}

.filters label,
.map-search label {
  display: grid;
  gap: 7px;
  color: var(--science-muted);
  font: 700 var(--brand-text-sm)/1.3 var(--brand-font-ui);
}

.filters :is(input, select),
.map-search input {
  width: 100%;
  min-height: 48px;
  padding: 10px 12px;
  border: 1px solid var(--science-line);
  border-radius: var(--brand-radius-sm);
  background: var(--science-bg);
  color: var(--science-ink);
}

.primary-button,
.taxonomy-filter {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  min-height: 48px;
  padding: 10px 16px;
  border: 1px solid var(--science-link);
  border-radius: var(--brand-radius-sm);
  background: var(--science-link);
  color: var(--science-bg);
  font: 700 var(--brand-text-sm)/1 var(--brand-font-ui);
  cursor: pointer;
}

.results-meta {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 24px;
  padding: 16px 0;
  border-top: 1px solid var(--science-line);
}

.results-meta p { margin: 0; color: var(--science-muted); }
.text-button { border: 0; background: transparent; cursor: pointer; }

.taxa-grid {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 1px;
  border: 1px solid var(--science-line);
  background: var(--science-line);
}

.taxon-card {
  min-height: 184px;
  padding: 22px;
  border: 0;
  background: var(--science-bg);
  color: var(--science-ink);
  text-align: left;
  cursor: pointer;
}

.taxon-card:hover { background: var(--science-surface-2); }
.taxon-card .rank { color: var(--science-moss); }
.taxon-card h3 { margin: 12px 0 3px; font: italic 700 1.3rem/1.2 var(--brand-font-display); }
.taxon-card p { margin: 4px 0; color: var(--science-muted); }
.taxon-card .lineage { font-size: .88rem; }

.empty-state {
  grid-column: 1 / -1;
  margin: 0;
  padding: 54px 24px;
  background: var(--science-bg);
  color: var(--science-muted);
  text-align: center;
}

.error { color: var(--science-danger) !important; }

.pagination {
  display: flex;
  justify-content: center;
  align-items: center;
  gap: 14px;
  margin-top: 24px;
}

.pagination button,
.copy-citation {
  min-height: 44px;
  padding: 8px 14px;
  border: 1px solid var(--science-line);
  border-radius: var(--brand-radius-sm);
  background: var(--science-bg);
  color: var(--science-ink);
  cursor: pointer;
}

.pagination button:disabled,
.copy-citation:disabled { opacity: .46; cursor: not-allowed; }
.pagination span { font-variant-numeric: tabular-nums; }

.detail-panel {
  position: fixed;
  inset: calc(var(--shell-height) + 16px) 16px 16px auto;
  z-index: 700;
  width: min(540px, calc(100% - 32px));
  overflow: auto;
  padding: clamp(24px, 4vw, 44px);
  border: 1px solid var(--science-line);
  background: var(--science-bg);
  box-shadow: var(--brand-shadow-3);
}

.detail-close {
  float: right;
  width: 44px;
  height: 44px;
  border: 1px solid var(--science-line);
  background: var(--science-surface);
  color: var(--science-ink);
  font-size: 1.4rem;
  cursor: pointer;
}

.authorship { color: var(--science-muted); font-style: italic; }
.taxonomy-grid { display: grid; grid-template-columns: 1fr 1fr; gap: 0 16px; margin: 24px 0; }
.detail-actions, .source-links { display: flex; flex-wrap: wrap; gap: 8px 18px; }

.source-note {
  display: grid;
  grid-template-columns: minmax(220px, .72fr) minmax(0, 1.28fr);
  gap: clamp(36px, 7vw, 96px);
  padding: clamp(64px, 9vw, 112px) 0;
}

.source-note p { color: var(--science-muted); }
.citation-block { margin-top: 24px; padding-top: 18px; border-top: 1px solid var(--science-line); }
.citation-block h3 { margin: 0 0 10px; font-family: var(--brand-font-display); }
.citation-text { overflow-wrap: anywhere; }

.coverage-banner {
  max-width: 880px;
  margin-top: 28px;
  padding: 18px 20px;
  border: 1px solid color-mix(in srgb, var(--science-accent) 55%, var(--science-line));
  background: color-mix(in srgb, var(--science-accent) 18%, var(--science-bg));
}

.coverage-banner strong { font-variant-numeric: tabular-nums; }
.coverage-banner span:last-child { display: block; color: var(--science-muted); }
.coverage-banner.unavailable { border-color: var(--science-line); background: var(--science-surface); }

.map-search {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 32px;
}

.map-search form div { display: flex; gap: 8px; }
.map-search-results { grid-column: 2; }

.search-result {
  display: block;
  width: 100%;
  min-height: 44px;
  padding: 10px 12px;
  border: 0;
  border-bottom: 1px solid var(--science-line);
  background: var(--science-surface-2);
  color: var(--science-ink);
  text-align: left;
  cursor: pointer;
}

.map-layout {
  display: grid;
  grid-template-columns: minmax(260px, .35fr) minmax(0, .65fr);
  min-height: 570px;
  margin-top: 16px;
  border: 1px solid var(--science-line);
  background: var(--science-surface);
}

.map-copy { padding: clamp(26px, 4vw, 48px); }
.map-canvas { min-height: 570px; background: #dfe5dd; }
.leaflet-container { font-family: var(--brand-font-body); }
.map-unavailable { display: grid; place-items: center; padding: 30px; background: var(--science-surface-2); color: var(--science-muted); text-align: center; }

.table-wrap { max-width: 100%; overflow: auto; margin-top: 20px; }
table { width: 100%; border-collapse: collapse; text-align: left; }
th, td { padding: 13px 12px; border-bottom: 1px solid var(--science-line); vertical-align: top; }
th { color: var(--science-muted); font: 700 var(--brand-text-xs)/1.3 var(--brand-font-ui); letter-spacing: .07em; text-transform: uppercase; }
tbody th { color: var(--science-ink); text-transform: none; letter-spacing: 0; }

/* Taxonomy is a snapshot-derived catalogue hierarchy, never a phylogeny. */
.taxonomy-scale-controls {
  display: flex;
  align-items: center;
  flex-wrap: wrap;
  gap: 8px;
  margin: 24px 0;
}

.taxonomy-scale-controls > span {
  margin-right: 4px;
  color: var(--science-muted);
  font: 700 var(--brand-text-xs)/1.3 var(--brand-font-ui);
  letter-spacing: .06em;
  text-transform: uppercase;
}

.taxonomy-scale {
  min-height: 44px;
  padding: 10px 14px;
  border: 1px solid var(--science-line);
  border-radius: var(--brand-radius-sm);
  border-color: var(--science-line);
  background: var(--science-bg);
  color: var(--science-ink);
  font: 700 var(--brand-text-sm)/1.2 var(--brand-font-ui);
  cursor: pointer;
}

.taxonomy-scale[aria-pressed="true"] {
  border-color: var(--science-link);
  background: var(--science-link);
  color: var(--science-bg);
}

.taxonomy-universe [hidden] { display: none !important; }
.taxonomy-status { color: var(--science-muted); }

.taxonomy-universe-layout {
  display: grid;
  grid-template-columns: minmax(0, 1.02fr) minmax(320px, .98fr);
  gap: clamp(28px, 5vw, 72px);
  align-items: start;
  margin-top: 28px;
}

.taxonomy-rings-figure { min-width: 0; margin: 0; }

.taxonomy-ring-canvas {
  position: relative;
  width: min(100%, 640px);
  aspect-ratio: 1;
  margin-inline: auto;
}

.taxonomy-ring-canvas svg { display: block; width: 100%; height: 100%; overflow: visible; }

.taxonomy-ring-track {
  fill: none;
  stroke: var(--science-line);
  stroke-width: 1;
}

.taxonomy-ring-segment {
  fill: none;
  stroke-linecap: butt;
  opacity: .9;
  transition: opacity var(--brand-duration-fast) var(--brand-ease-out-quad),
    stroke-width var(--brand-duration-fast) var(--brand-ease-out-quad);
}

.taxonomy-ring-segment.is-boundary { stroke-dasharray: 10 5; }
.taxonomy-ring-segment.is-muted { opacity: .16; }
.taxonomy-ring-segment.is-active { opacity: 1; }
.taxonomy-ring-segment--phylum.is-active { stroke-width: 62px !important; }
.taxonomy-ring-segment--order.is-active { stroke-width: 42px !important; }

.taxonomy-ring-readout {
  position: absolute;
  inset: 50% auto auto 50%;
  display: grid;
  width: 32%;
  min-width: 120px;
  translate: -50% -50%;
  text-align: center;
}

.taxonomy-ring-readout span {
  color: var(--science-muted);
  font: 700 var(--brand-text-xs)/1.3 var(--brand-font-ui);
  letter-spacing: .04em;
}

.taxonomy-ring-readout strong {
  margin: 4px 0;
  color: var(--science-ink);
  font: 700 clamp(1.7rem, 4.5vw, 3.2rem)/.95 var(--brand-font-display);
  font-variant-numeric: tabular-nums;
}

.taxonomy-rings-figure figcaption {
  max-width: 60ch;
  margin: 16px auto 0;
  color: var(--science-muted);
  font-size: var(--brand-text-sm);
}

.taxonomy-ring-legend {
  display: flex;
  flex-wrap: wrap;
  gap: 8px 18px;
  margin: 16px 0 0;
  padding: 16px 0 0;
  border-top: 1px solid var(--science-line);
  color: var(--science-muted);
  font-size: var(--brand-text-xs);
  list-style: none;
}

.taxonomy-ring-legend li { display: flex; align-items: center; gap: 7px; }
.legend-line { display: inline-block; width: 26px; border-top: 5px solid var(--brand-indigo-600); }
.legend-line--order { border-top-width: 3px; border-color: var(--brand-ochre-700); }
.legend-line--boundary { border-top: 3px dashed var(--science-muted); }

.taxonomy-hierarchy-list { display: grid; gap: 32px; min-width: 0; }
.taxonomy-hierarchy-list section { min-width: 0; }

.taxonomy-list-heading {
  display: flex;
  align-items: baseline;
  justify-content: space-between;
  gap: 16px;
  padding-bottom: 10px;
  border-bottom: 1px solid var(--science-line);
}

.taxonomy-list-heading h3 { margin: 0; font: 700 1.25rem/1.15 var(--brand-font-display); }
.taxonomy-list-heading span,
.taxonomy-selection { color: var(--science-muted); font-size: var(--brand-text-sm); }
.taxonomy-selection { margin: 12px 0; }

.taxonomy-node-list {
  display: grid;
  margin: 0;
  padding: 0;
  list-style: none;
}

.taxonomy-node-list--orders { max-height: 430px; overflow: auto; }

.taxonomy-node {
  position: relative;
  display: grid;
  grid-template-columns: minmax(0, 1fr) auto;
  min-width: 0;
  border-bottom: 1px solid var(--science-line);
}

.taxonomy-node.is-unassigned::before {
  position: absolute;
  inset: 0;
  z-index: 0;
  background: repeating-linear-gradient(135deg, transparent 0 8px, color-mix(in srgb, var(--science-muted) 9%, transparent) 8px 11px);
  content: '';
  pointer-events: none;
}

.taxonomy-node.is-boundary { border-left: 2px dashed var(--science-muted); }

.taxonomy-node-control,
.taxonomy-node-link {
  position: relative;
  z-index: 1;
}

.taxonomy-node-control {
  display: flex;
  min-width: 0;
  min-height: 48px;
  align-items: center;
  justify-content: space-between;
  gap: 14px;
  padding: 9px 10px;
  border: 0;
  background: transparent;
  color: var(--science-ink);
  text-align: left;
}

button.taxonomy-node-control { cursor: pointer; }
button.taxonomy-node-control:hover,
button.taxonomy-node-control[aria-pressed="true"] { background: color-mix(in srgb, var(--science-link) 9%, transparent); }
button.taxonomy-node-control[aria-pressed="true"] strong { text-decoration: underline; text-decoration-thickness: 2px; text-underline-offset: .28em; }

.taxonomy-node-control strong { overflow: hidden; font: 700 var(--brand-text-sm)/1.25 var(--brand-font-body); text-overflow: ellipsis; }
.taxonomy-node-control span { flex: none; color: var(--science-muted); font-size: var(--brand-text-xs); font-variant-numeric: tabular-nums; }

.taxonomy-node-link {
  display: inline-flex;
  min-height: 48px;
  align-items: center;
  padding: 8px 10px;
  color: var(--science-link);
  font: 700 var(--brand-text-xs)/1.2 var(--brand-font-ui);
  text-underline-offset: .2em;
}

.taxonomy-live {
  position: absolute;
  width: 1px;
  height: 1px;
  margin: -1px;
  padding: 0;
  overflow: hidden;
  border: 0;
  clip: rect(0 0 0 0);
  white-space: nowrap;
}

.family-universe .science-section-heading { align-items: start; }
.family-universe [hidden] { display: none !important; }

.family-load-status,
.family-results-status {
  margin: 24px 0 0;
  color: var(--science-muted);
}

.family-snapshot {
  display: grid;
  grid-template-columns: 1.3fr 1fr 1fr 1fr;
  gap: 1px;
  margin: 24px 0 0;
  border: 1px solid var(--science-line);
  background: var(--science-line);
}

.family-snapshot div {
  min-width: 0;
  padding: 16px;
  background: var(--science-bg);
}

.family-snapshot dt {
  font: 700 var(--brand-text-xs)/1.3 var(--brand-font-ui);
  letter-spacing: .055em;
  text-transform: uppercase;
}

.family-snapshot dd {
  margin-top: 7px;
  font-variant-numeric: tabular-nums;
}

.family-controls {
  display: grid;
  grid-template-columns: minmax(240px, 1fr) minmax(210px, .38fr) auto;
  gap: 16px;
  align-items: end;
  margin-top: 28px;
}

.family-controls label {
  display: grid;
  gap: 7px;
  color: var(--science-muted);
  font: 700 var(--brand-text-sm)/1.3 var(--brand-font-ui);
}

.family-controls :is(input, select) {
  width: 100%;
  min-height: 48px;
  padding: 10px 12px;
  border: 1px solid var(--science-line);
  border-radius: var(--brand-radius-sm);
  background: var(--science-bg);
  color: var(--science-ink);
}

.family-toggle {
  min-height: 48px;
  padding: 10px 16px;
  border: 1px solid var(--science-link);
  border-radius: var(--brand-radius-sm);
  background: transparent;
  color: var(--science-link);
  font: 700 var(--brand-text-sm)/1 var(--brand-font-ui);
  cursor: pointer;
}

.family-table-wrap {
  max-height: 680px;
  margin-top: 16px;
  overflow: auto;
  border: 1px solid var(--science-line);
  background: var(--science-bg);
}

.family-table caption {
  padding: 14px 16px;
  color: var(--science-muted);
  text-align: left;
}

.family-table thead {
  position: sticky;
  z-index: 1;
  top: 0;
  background: var(--science-surface-2);
}

.family-table tbody th {
  font-family: var(--brand-font-body);
  font-style: italic;
}

.family-count {
  font-variant-numeric: tabular-nums;
  text-align: right;
}

.family-catalog-link {
  display: inline-flex;
  align-items: center;
  min-height: 44px;
  color: var(--science-link);
  font: 700 var(--brand-text-sm)/1.2 var(--brand-font-ui);
  text-underline-offset: .2em;
}

.family-empty { padding: 36px 16px; color: var(--science-muted); text-align: center; }

.family-provenance,
.family-unavailable {
  margin-top: 20px;
  padding: 18px 20px;
  border: 1px solid var(--science-line);
  background: var(--science-surface-2);
  color: var(--science-muted);
}

.family-provenance p,
.family-unavailable p { margin: 0; }
.family-provenance p + p { margin-top: 8px; }
.family-provenance a { color: var(--science-link); }
.family-unavailable h3 { margin: 0 0 8px; color: var(--science-ink); font: 700 1.2rem var(--brand-font-display); }

.atlas-grid {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 16px;
  margin-top: 24px;
}

.atlas-entry { margin: 0; border: 1px solid var(--science-line); background: var(--science-bg); }
.atlas-entry img { display: block; width: 100%; aspect-ratio: 4 / 3; object-fit: cover; background: var(--science-surface-2); }
.atlas-entry figcaption { padding: 16px; }
.atlas-entry h3 { margin: 0 0 5px; font: italic 700 1.2rem/1.2 var(--brand-font-display); }
.atlas-entry p { margin: 7px 0; color: var(--science-muted); font-size: .9rem; }

.glossary-grid,
.capability-grid {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 1px;
  margin-top: 24px;
  border: 1px solid var(--science-line);
  background: var(--science-line);
}

.glossary-grid div,
.capability-card { padding: 22px; background: var(--science-bg); }
.glossary-grid dt { color: var(--science-ink); font: 700 1.05rem var(--brand-font-display); }
.glossary-grid dd { margin-top: 8px; color: var(--science-muted); font-weight: 400; }

/* Data is an inventory: statuses and provenance take precedence over decor. */
.capability-card h3 { margin: 0; font: 700 1.2rem var(--brand-font-display); }
.capability-card p { color: var(--science-muted); }
.capability-state--available { color: var(--science-moss); }
.capability-state--bounded { color: var(--science-link); }
.capability-state--not-public { color: var(--science-muted); }

.code-sample {
  margin: 24px 0 0;
  padding: 20px;
  overflow: auto;
  border: 1px solid var(--science-line);
  background: var(--brand-neutral-950);
  color: var(--brand-cream-100);
  font: 400 .88rem/1.65 var(--brand-font-mono);
}

.science-note {
  margin-top: 20px;
  padding: 18px 20px;
  border: 1px solid var(--science-line);
  background: var(--science-surface-2);
  color: var(--science-muted);
}

@media (max-width: 900px) {
  .catalog-hero,
  .science-hero--split,
  .map-search,
  .map-layout,
  .source-note { grid-template-columns: 1fr; }

  .snapshot-context .snapshot-meta { grid-template-columns: repeat(2, minmax(0, 1fr)); }
  .filters { grid-template-columns: 1fr 1fr; }
  .filters .primary-button { grid-column: 1 / -1; }
  .taxa-grid,
  .atlas-grid,
  .glossary-grid,
  .capability-grid,
  .family-snapshot { grid-template-columns: repeat(2, minmax(0, 1fr)); }
  .family-controls { grid-template-columns: 1fr 1fr; }
  .family-toggle { grid-column: 1 / -1; }
  .taxonomy-universe-layout { grid-template-columns: 1fr; }
  .taxonomy-ring-canvas { width: min(100%, 560px); }
  .map-search-results { grid-column: 1; }
  .map-copy { min-height: 250px; }
}

@media (max-width: 600px) {
  body[data-page="taxonomy"] main,
  body[data-page="catalog"] main,
  body[data-page="map"] main,
  body[data-page="data"] main { width: min(calc(100% - 24px), var(--science-max)); }

  .catalog-hero,
  .map-hero,
  .science-hero { padding-top: 44px; }

  .catalog-hero h1,
  .map-hero h1,
  .science-hero h1 { font-size: clamp(2.35rem, 13vw, 3.35rem); }

  .catalog-shell,
  .map-search,
  .table-section,
  .taxonomy-workbench,
  .science-panel { padding: 20px; }

  .section-heading,
  .science-section-heading,
  .results-meta { align-items: flex-start; flex-direction: column; gap: 12px; }
  .filters,
  .taxa-grid,
  .atlas-grid,
  .glossary-grid,
  .capability-grid,
  .snapshot-meta,
  .snapshot-context .snapshot-meta,
  .taxonomy-grid { grid-template-columns: 1fr; }
  .family-snapshot,
  .family-controls { grid-template-columns: 1fr; }
  .family-toggle { grid-column: auto; }
  .map-search form div { align-items: stretch; flex-direction: column; }
  .map-canvas { min-height: 400px; }
  .pagination { flex-wrap: wrap; }
  th, td { min-width: 132px; }
  .taxonomy-node { grid-template-columns: minmax(0, 1fr) auto; }
  .taxonomy-node-control { align-items: flex-start; flex-direction: column; gap: 3px; }
  .taxonomy-node-link { padding-inline: 7px; }
  .taxonomy-ring-readout { min-width: 96px; }
}

@media print {
  .taxonomy-scale-controls { display: none; }
  .taxonomy-universe-layout { grid-template-columns: .9fr 1.1fr; gap: 24px; }
  .taxonomy-node-list--orders { max-height: none; overflow: visible; }
  .taxonomy-node-link { display: none; }
  .taxonomy-ring-segment { transition: none; }
}

@media (prefers-reduced-motion: reduce) {
  html { scroll-behavior: auto; }
  *, *::before, *::after {
    scroll-behavior: auto !important;
    animation-duration: .01ms !important;
    transition-duration: .01ms !important;
  }
}